1. Robert W. Holley Winner Of The 1968 Nobel Prize In Medicine robert W. holley, a nobel Prize Laureate in Physiology and Medicine, at the nobelPrize Internet Archive. robert W. holley. 1968 nobel Laureate in Medicine http://almaz.com/nobel/medicine/1968a.html

R OBERT W H OLLEY 1968 Nobel Laureate in Medicine for their interpretation of the genetic code and its function in protein synthesis. Background Born: 1922 Residence: U.S.A. Affiliation: Cornell University, Ithaca, NY Featured Internet Links Prize co-recipient: Har Gobind Khorana Prize co-recipient: Marshall W. Nirenberg Search WWW Search The Nobel Prize Internet Archive Links added by Nobel Internet Archive visitors Back to The Nobel Prize Internet Archive Literature Peace Chemistry ... Medicine We always welcome your feedback and comments

2. Index Of Nobel Laureates In Medicine ALPHABETICAL LISTING OF nobel PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE.Name, Year Awarded. Hodgkin, Sir Alan Lloyd, 1963. holley, robert W. 1968. http://almaz.com/nobel/medicine/alpha.html

ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE Name Year Awarded Adrian, Lord Edgar Douglas Arber, Werner Axelrod, Julius Baltimore, David ... Medicine We always welcome your feedback and comments

3. Medicine 1968 Provides details of the 1968 nobel prize awarded for their interpretation of the genetic code and Category Science Biology Genetics History People......The nobel Prize in Physiology or Medicine 1968. for their interpretation of thegenetic code and its function in protein synthesis . robert W. holley, Har Gobind http://www.nobel.se/medicine/laureates/1968/

The Nobel Prize in Physiology or Medicine 1968 "for their interpretation of the genetic code and its function in protein synthesis" Robert W. Holley Har Gobind Khorana Marshall W. Nirenberg 1/3 of the prize 1/3 of the prize 1/3 of the prize USA USA USA Cornell University Ithaca, NY, USA University of Wisconsin Madison, WI, USA National Institutes of Health Bethesda, MD, USA b. 1922 d. 1993 b. 1922 (in Raipur, India) b. 1927 The Nobel Prize in Physiology or Medicine 1968 Presentation Speech Robert W. Holley Biography ... Other Resources The 1968 Prize in: Physics Chemistry Physiology or Medicine Literature ... Peace Find a Laureate: Last modified June 16, 2000 The Official Web Site of The Nobel Foundation

4. Robert W. Holley - Biography robert W. holley was born in Urbana, Illinois, on January 28th holley's training asa chemist did not alter his basic was for this work that the nobel Prize was http://www.nobel.se/medicine/laureates/1968/holley-bio.html

Robert W. Holley was born in Urbana, Illinois, on January 28th, 1922, one of four sons of Charles and Viola Holley. His parents were both educators. He attended public schools in Illinois, California and Idaho, and graduated from Urbana High School in 1938. He studied chemistry at the University of Illinois and received his B. A. degree in 1942. Graduate work was at Cornell University , where the Ph.D. degree in organic chemistry, with Professor Alfred T. Blomquist, was awarded in 1947. Graduate work was interrupted during the war. He spent two years, 1944-1946, with Professor Vincent du Vigneaud at Cornell University Medical College , where he participated in the first chemical synthesis of penicillin. After completing the Ph. D. degree, Holley spent 1947-1948 as an American Chemical Society Postdoctoral Fellow with Professor Carl M. Stevens at Washington State University . He then returned to Cornell University as Assistant Professor of Organic Chemistry at the Geneva Experiment Station in 1948. He was Associate Professor there from 1950-1957. During a sabbatical year, 1955-1956, he was a

5. Holley, Robert William holley, robert William. (b. Jan. 28, 1922, Urbana, Ill., USd. Feb. 11, 1993, LosGatos, Calif.), American biochemist who shared the nobel Prize in Physiology http://www.britannica.com/nobel/micro/274_63.html

Holley, Robert William (b. Jan. 28, 1922, Urbana, Ill., U.S.d. Feb. 11, 1993, Los Gatos, Calif.), American biochemist who shared the Nobel Prize in Physiology or Medicine in 1968 with Marshall Warren Nirenberg and Har Gobind Khorana . Their research helped explain how the genetic code controls the synthesis of proteins. Holley obtained his Ph.D. in organic chemistry from Cornell University, Ithaca, N.Y., in 1947. He investigated a variety of biochemical questions at the state and federal agricultural experiment stations at Cornell (1948-64). He began his research on RNA after spending a year studying with James F. Bonner at the California Institute of Technology (1955-56). By 1960 Holley and others had shown that small molecules of ribonucleic acids, called transfer RNAs, were involved in the assembly of amino acids into proteins. Holley and his collaborators developed techniques to separate the different transfer RNAs from the mixture in the cell. By 1965 he had determined the composition of the transfer RNA that incorporates the amino acid alanine into protein molecules. This featthe first determination of the sequence of nucleotides in a nucleic acidrequired digesting the molecule with enzymes, identifying the pieces, then figuring out how they fit together. It has since been shown that all transfer RNAs have similar structures. In 1968 Holley became a resident fellow at the Salk Institute for Biological Studies in La Jolla, Calif. He also became an adjunct professor at the University of California, San Diego, in the following year.

6. Nobel Prize Winners For 1961-1970 physics, Alvarez, Luis W. US, work with elementary particles, discovery of resonancestates, physiology/medicine, holley, robert William, US, deciphering of the http://www.britannica.com/nobel/1961_70.html

Year Category Article Country* Achievement Literary Area chemistry Calvin, Melvin U.S. study of chemical steps that take place during photosynthesis literature Andric, Ivo Yugoslavia novelist peace Sweden physics Hofstadter, Robert U.S. determination of shape and size of atomic nucleons physics West Germany physiology/medicine U.S. functions of the inner ear chemistry Kendrew, Sir John Cowdery U.K. determination of the structure of hemoproteins chemistry Perutz, Max Ferdinand U.K. determination of the structure of hemoproteins literature Steinbeck, John U.S. novelist peace Pauling, Linus U.S. physics Landau, Lev Davidovich U.S.S.R. contributions to the understanding of condensed states of matter physiology/medicine Crick, Francis Harry Compton U.K. discoveries concerning the molecular structure of DNA physiology/medicine Watson, James Dewey U.S. discoveries concerning the molecular structure of DNA physiology/medicine Wilkins, Maurice U.K. discoveries concerning the molecular structure of DNA chemistry Natta, Giulio

7. ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE FRONTIERS IN BIOSCIENCE; ALPHABETICAL LISTING OF nobel PRIZE LAUREATESIN PHYSIOLOGY AND MEDICINE, Name, Year Awarded. holley, robert W. 1968. http://www.bioscience.org/urllists/nobelm.htm

FRONTIERS IN BIOSCIENCE; ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE Name Year Awarded Adrian, Lord Edgar Douglas Arber, Werner Axelrod, Julius Baltimore, David ... Zinkernagel, Rolf M.

8. ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY Zsigmondy, Richard Adolf, 1925. ALPHABETICAL LISTING OF nobel PRIZE LAUREATES INPHYSIOLOGY AND MEDICINE. Hodgkin, Sir Alan Lloyd, 1963. holley, robert W. 1968. http://www.bioscience.org/urllists/nobelc.htm

FRONTIERS IN BIOSCIENCE; ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY, PHYSIOLOGY AND MEDICINE ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY Name Year Awarded Alder, Kurt Altman, Sidney Anfinsen, Christian B. Arrhenius, Svante August ... Zsigmondy, Richard Adolf ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE Name Year Awarded Adrian, Lord Edgar Douglas Arber, Werner Axelrod, Julius Baltimore, David ... Zinkernagel, Rolf M. Source: The Nobel Prize Internet Archive

9. Premios Nobel De Medicina Premios nobel de Medicina. Año, Tema, Ganador. 1901, Behring, Emil AdolfVon. 1968, holley, robert W.; Khorana, Har Gobind; Nirenberg, Marshall W. http://fai.unne.edu.ar/biologia/nobeles/nobelmed.htm

Premios Nobel de Medicina Tema Ganador Behring, Emil Adolf Von Ross, Sir Ronald Finsen, Niels Ryberg Pavlov, Ivan Petrovich Koch, Robert Cajal, Santiago Ramon Y.; Golgi, Camillo Laveran, Charles Louis Alphonse Ehrlich, Paul; Metchnikoff, Ilya Ilyich Kocher, Emil Theodor Kossel, Albrecht Gullstrand, Allvar Carrel, Alexis Richet, Charles Robert Barany, Robert Bordet, Jules Krogh, Schack August Steenberger Hill, Sir Archibald Vivian; Meyerhof, Otto Fritz; Banting, Sir Frederick Grant; Macleod, John James Richard; Einthoven, Willem; Fibiger, Johannes Andreas Grib Wagner-Jauregg, Julius Nicolle, Charles Jules Henri Eijkman, Christiaan; Hopkins, Sir Frederick Gowland Landsteiner, Karl Warburg, Otto Heinrich Adrian, Lord Edgar Douglas; Sherrington, Sir Charles Scott Morgan, Thomas Hunt Minot, George Richards; Murphy, William Parry; Whipple, George Hoyt Spemann, Hans Dale, Sir Henry Hallett; Loewi, Otto Nagyrapolt, Albert Szent-Gyorgyi Von Heymans, Corneille Jean Francois Domagk, Gerhard Dam, Henrik Carl Peter; Doisy, Edward Adelbert Erlanger, Joseph; Gasser, Herbert Spencer

10. Nobel Laureates - 7. Lectures And Nobel Laureates - NIH 1999 Almanac Content nobel Laureates. Laureate, Field, Year, Supporting Institute(s). robert W. holley,USA (shared with HG Khorana and MW Nirenberg, USA), .do, 1968, NIGMS, NCI. http://www.nih.gov/about/almanac/1999/lectures/nobel.html

NIH 1999 Almanac Lectures and Nobel Laureates Nobel Laureates Laureate Field Year Supporting Institute(s) Robert Furchgott, U.S.A. (shared with L. Ignarro and F. Murad, U.S.A.) Physiology or medicine NIGMS, NHLBI, NINDS Louis Ignarro, U.S.A. (shared with F. Murad, and R. Furchgott. U.S.A.) ......do NHLBI, NIAMS, NICHD Ferid Murad, U.S.A. (shared with L. Ignarro, and R. Furchgott. U.S.A.) ......do NIAMS, NIGMS, NHLBI, NIDDK Paul D. Boyer, U.S.A. (shared with J.C. Skou) Chemistry NIGMS, NIDDK Jens C. Skou, Denmark (shared with P.D. Boyer) ......do NINDS Stanley B. Prusiner, U.S.A. Phyisology or medicine NINDS, NIA, NCRR, NIGMS Edward B. Lewis, U.S.A. (shared with C. Nusslein-Volhard, Germany, and E.F. Wieschaus, U.S.A.) Physiology or medicine NICHD, NIGMS Eric F. Wieschaus, U.S.A. (shared with E.B. Lewis, U.S.A., and C. Nusslein-Volhard, Germany) ......do NICHD Alfred G. Gilman, U.S.A. (shared with M. Rodbell, U.S.A.) .....do NIGMS, NINDS Martin Rodbell, U.S.A. (shared with A.G. Gilman, U.S.A.) ......do NIEHS, NIDDK George A. Olah, U.S.A. Chemistry NCI, NIGMS

11. NIH: About: NIH Almanac: Nobel Laureates Read about the NIH Scientists who have won nobel prizes. robert W. holley, USA(shared with HG Khorana and MW Nirenberg, USA), , 1968, NIGMS, NCI. http://www.nih.gov/about/almanac/nobel/

Home About NIH NIH Almanac About the Almanac ... Past Issues Nobel Laureates Read about the NIH Scientists who have won Nobel prizes. Laureate Field Year Supporting NIH Institute(s) Leland H. Hartwell, U.S.A. (shared with P.M. Nurse and R.T. Hunt, U.K.) Physiology or medicine NIGMS, NCI, NCRR K. Barry Sharpless, U.S.A. (shared with W.S. Knowles, U.S.A. and R. Noyori, Japan) Chemistry NIGMS, NHLBI Paul Greengard, U.S.A. (shared with E. Kandel, U.S.A. and A. Carlsson, Sweden) Physiology or medicine NIMH, NIA, NIDA, NINDS, NIAAA, NHLBI, NIAMS Erik R. Kandel, U.S.A. (shared with P. Greengard, U.S.A. and A. Carlsson, Sweden) " NIMH, NIGMS, NINDS, NCRR James J. Heckman, U.S.A. (shared with D. McFadden, U.S.A.) Economic sciences NICHD, NIMH Daniel L. McFadden, U.S.A. (shared with J. Heckman, U.S.A.) " NIA Günter Blobel, U.S.A. Physiology or medicine NIGMS, NCI Robert Furchgott, U.S.A. (shared with L. Ignarro and F. Murad, U.S.A.) Physiology or medicine NIGMS, NHLBI, NINDS

12. Sigma Xi: The Scientific Research Society: Nobel Laureates About Sigma Xi » Overview » nobel Laureates 1966 Francis Peyton Rous 1967 HaldanKeffer Hartline 1967 George Wald 1968 robert W. holley 1968 Marshall W http://www.sigmaxi.org/about/overview/nobel.shtml

Overview Leadership Organization News ... Contact Us About: Overview Leadership Organization News Support Sigma Xi ... Contact Us Search: Site Map Home Overview Physics 1907 Albert Michelson 1921 Albert Einstein 1923 Robert A. Millikan 1925 James Franck 1927 Arthur H. Compton 1936 Carl D. Anderson 1937 Clinton J. Davisson 1938 Enrico Fermi 1939 Ernest O. Lawrence 1943 Otto Stern 1944 Isidor Isaac Rabi 1945 Wolfgang Pauli 1946 Percy Williams Bridgman 1952 Felix Bloch 1952 Edward M. Purcell 1955 Polykarp Kusch 1955 Willis E. Lamb, Jr. 1956 John Bardeen 1956 Walter H. Brattain 1956 William Shockley 1957 Chen Ning Yang 1958 Igor Y. Tamm 1959 Owen Chamberlain 1959 Emilio G. Segre 1960 Donald A. Glaser 1961 Robert Hofstadter 1963 Eugene P. Wigner

13. Nobel Prize In Physiology Or Medicine - Wikipedia Source http//www.nobel.se/medicine/laureates/index.html. Huggins 1967 Ragnar Granit,Haldan Keffer Hartline, George Wald 1968 robert W. holley, Har Gobind http://www.wikipedia.org/wiki/Nobel_Prize/Physiology_or_medicine

Main Page Recent changes Edit this page Page history Special pages Set my user preferences My watchlist Recently updated pages Upload image files Image list Registered users Site statistics Random article Orphaned articles Orphaned images Popular articles Most wanted articles Short articles Long articles Newly created articles Interlanguage links All pages by title Blocked IP addresses Maintenance page External book sources Printable version Talk Log in Help Other languages: Deutsch Dansk Nederlands Polski Nobel Prize in Physiology or Medicine (Redirected from Nobel Prize/Physiology or medicine Emil Adolf von Behring Ronald Ross Niels Ryberg Finsen ... Christiaan Eijkman , Sir Frederick Gowland Hopkins Karl Landsteiner Otto Heinrich Warburg Sir Charles Scott Sherrington Edgar Douglas Adrian Thomas Hunt Morgan George Hoyt Whipple ... Hans Spemann Sir Henry Hallett Dale Otto Loewi Albert von Szent-Györgyi Nagyrapolt Corneille Jean François Heymans ... Herbert Spencer Gasser Sir Alexander Fleming Ernst Boris Chain , Sir Howard Walter Florey Hermann Joseph Muller Carl Ferdinand Cori Gerty Theresa , née Radnitz Cori, Bernardo Alberto Houssay Paul Hermann Müller Walter Rudolf Hess Antonio Caetano De Abreu Freire Egas Moniz ... Dickinson W. Richards

14. Cornell Nobel Laureates robert W. holley, Professor and Department Chair in Biochemistry (194864), 1968Prize in Medicine; Updated 2/8/00. Return to nobel Prize release. http://www.news.cornell.edu/science/Oct96/Nobel_Laureates_at_CU.html

Nobel Laureates Affiliated with Cornell University Compiled by the Cornell News Service, 1995 Twenty-nine Nobel Prize winners have been affiliated with Cornell University as alumni or faculty members. Current Faculty Members Hans Bethe, the John Wendell Anderson Professor of Physics Emeritus, 1967 Prize in Physics Roald Hoffmann, the Frank H.T. Rhodes Professor in Humane Letters, 1981 Prize in Chemistry David M. Lee, professor of physics, 1996 Nobel Laureate in Physics Robert C. Richardson, the Floyd R. Newman Professor of Physics, 1996 Nobel Laureate in Physics Alumni George W. Beadle, '31 Ph.D. Genetics, 1958 Prize in Physiology and Medicine (also was professor of agriculture) Pearl Buck, '25 M.A., 1938 Prize in Literature Robert W. Fogel, '48 A.B. Economics, 1993 Prize in Economics Sheldon Glashow, '54 B.S. Physics, 1979 Prize in Physics Barbara McClintock, '23 B.S. Botany, '25 M.A., '27 Ph.D. Plant Genetics, 1983 Prize in Medicine Toni Morrison, '55 M.S. English, 1993 Prize in Literature John R. Mott, '88 B.S. Philosophy, 1946 Peace Prize Herman J. Muller, graduate study 1911-12, 1946 Prize for Medicine and Physiology

15. The Lasker Foundation | Former Award Winners, Basic Medical Research 1965, Obitu robert W. holley, a biologist who won the nobel Prize for unravelingthe genetic code of ribonucleic acid, died Thursday. He was 71. http://www.laskerfoundation.org/awards/obits/holleyobit.shtml

Lasker Awards Jury Members This Year's Winners Former Winners Library of Laureates Browse the Library for former winners of the Albert Lasker Basic Medical Research Award Roll your mouse over the left oval Nominations Open call to... Nominate a Scientist Former Winners, 1965 Albert Lasker Award for Basic Medical Research Robert W. Holley Obituary The following is an excerpt of an obituary for Dr. Holley published by the New York Times on Sunday, February 14, 1993. Robert W. Holley, a biologist who won the Nobel Prize for unraveling the genetic code of ribonucleic acid, died Thursday. He was 71. Dr. Holley was the first to unravel the internal structure in a strand of RNA, which helps determine what form and role each cell takes in a larger organism. Under the RNA's direction, building blocks of proteins become different parts of living things. The scientific structure he analyzed was alanine transfer RNA. His later research at Salk focused on the factors that stimulate or inhibit the growth of cells in mammals, and studies on the factors controlling the timing of cell division.

16. The Lasker Foundation | Lasker Awards And The Nobel Year of. Basic Award Winner, Lasker, nobel. George Wells Beadle, 1950, 1958. AlfredD. Hershey, 1958, 1969. robert W. holley, 1965, 1968. Eric R. Kandel, 1983, 2000. http://www.laskerfoundation.org/awards/n_vs_l.html

Lasker Awards Jury Members This Year's Winners Former Winners ... Award History View Video Interviews Of select Lasker Award winners... Lasker Luminaries The Awards Society Find out how you can participate in our deferred giving program... The Awards Society Lessons Learned The making of a premier Awards Program. Find out how in... Lessons Learned Nominations Open call to... Nominate a Scientist Lasker Awards and the Nobel The Lasker Medical Science Awards in basic research, clinical research, special achievement and public service, which have been bestowed since l945, provide a chronicle of the progress of biomedical research over the last half-century. The Lasker Foundation is proud that many of the amazing discoveries and achievements of Lasker Award winners are recognized, in addition, by the prestigious Nobel Prize. As of 2002, sixty-six Lasker winners have gone on to win the Nobel. The following statistics are of interest: 47.5% of the Basic Lasker Winners go on to win the Nobel

17. Premio Nobel De Medicina - Wikipedia Translate this page Ver enlace http//www.nobel.se/medicine/laureates/index.html. 1969 Max Delbrück,Alfred D. Hershey, Salvador E. Luria 1968 robert W. holley, Har Gobind http://es.wikipedia.org/wiki/Premio_Nobel/Medicina

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18. Prix Nobel De Physiologie Ou Médecine - Wikipedia Les prix nobel de physiologie et médecine. 1901 Emil Adolf von Behring. 1902 RonaldRoss. 1968 robert W. holley, Har Gobind Khorana, Marshall W. Nirenberg. http://fr.wikipedia.org/wiki/Prix_Nobel_de_Physiologie_ou_Médecine

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19. Nobel For Medicine: All Laureates E. Luria 1968 robert W. holley, Har Gobind Frédéric Cournand, Werner Forssmann,Dickinson W. Richards 1955 Axel Santiago Ramón y Cajal 1905 robert Koch 1904 http://www.popular-science.net/nobel/med-list.html

NOBEL Read More on the Nobel Prize: Nobel Minds 2001 Awards John Nash Women Nobel Prize ... Prize Amounts MINI ALMANAC Calendar Moon phase Popular Science Highlights: IG Nobel 2002 The invention of :-) West Nile Virus Asteroid Impact? ... Book: Russell Read also: Nobel Prize Women in Science : Their Lives, Struggles, and Momentous Discoveries by Sharon Bertsch McGrayne THE NOBEL PRIZE FOR MEDICINE: ALL WINNERS 2001 Leland H. Hartwell, R. Timothy Hunt, Paul M. Nurse 2000 Arvid Carlsson, Paul Greengard, Eric R. Kandel 1999 Günter Blobel 1998 Robert F. Furchgott, Louis J. Ignarro, Ferid Murad 1997 Stanley B. Prusiner 1996 Peter C. Doherty, Rolf M. Zinkernagel 1995 Edward B. Lewis, Christiane Nüsslein-Volhard, Eric F. Wieschaus 1994 Alfred G. Gilman, Martin Rodbell 1993 Richard J. Roberts, Phillip A. Sharp 1992 Edmond H. Fischer, Edwin G. Krebs

20. ŵ ±´ ¶û ½± Ïû Ï¢ robert W. holley Shared the nobel Prize in Physiology or Medicine 1968 With Har GobindKhorana (1922 , USA) and Marshall W. Nirenberg (1927- , USA) for their http://www.networkchinese.com/chineseprof/region/other/uc_camtus/sandiego/holley

Robert W. Holley Robert W. Holley Shared the Nobel Prize in Physiology or Medicine 1968 With Har Gobind Khorana (1922- , USA) and Marshall W. Nirenberg (1927- , USA) "for their interpretation of the genetic code and its function in protein synthesis" ºÉ À×£¬ À ¹ú ·Ö ×Ó Éú Îï ѧ ¼Ò£¬ Òò ÆÆ ½â ÒÅ ´« Ü Âë ²¢ ½â ÊÍ Æä µ° °× ºÏ ³É ¹¦ ÄÜ£¬ ¶ø Óë Áí Á½ λ À ¹ú ¿Æ ѧ ¼Ò »ô À­ ÄÇ (Har Gobind Khorana) ºÍ Äá Â× ²® (Marshall W. Nirenberg) ¹² »ñ 1968 Äê ŵ ±´ ¶û Ò½ ѧ ½±¡£ www.nobel.se/medicine/laureates/1968/holley-bio.html

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